Figured it out... this is so dumb!
So, turns out that there are two quests in that same area that each involve a jawbone with the exact same image in the inventory.
Once I realised that the "dusty old jawbone" was a nodrop and the other items were droppable, I googled it & found out that it was for a completely different quest. I think I got it when I accidentally went inside the bear cave. After I went back to the river, I looted a "barbarian jawbone", which let me combine the box and turn in the quest, and I put the other jawbone in the bank in case I need it later.
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